Name                          position in the lab                       position after

Jin tang                     ph.d. student                                 research specialist, UCSF

Dr. Wenyan Xu          project scientist                          Analytical chemist, Eurofins Lancaster Laboratories


Dr. Chaoxing Liu      postdoc                                         Associate Professor, sun yat-sen uniVersity


Ching-Hsin Yang (2020-2022), MS student; Phd student, Academia Sinica

Anshu Joshi (2018-2020), MS student;

daniel oppong (2017-2019), Ms student; phd student, wayne state univ

Nana morehouse (2017-2019), MS student; Scientist, Oxford Natural Laboratories

maya tree (2017-2019), laboratory technician; central michigan univ

Riley boyd (2017-2018), visiting student/technician; Md student, northwestern univ

Faris samkari (2018-2019), BS student; MS student, kaust

madison tarnowski (summer 2017), summer visiting student, univ of michigan

Alan Huang (summer 2017), high school student; johns hopkins univ

OLiver Yockey (summer 2015 --- Summer 2017), BS student; md student, central michigan univ

Ashley Keating (Summer 2016 --- Spring 2017), BS student; MD student, Michigan state univ

Daniel kool (fall 2013 --- summer 2015), MS student; PhD student, Iowa State Univ

Tianzuo Xu (spring 2015 --- spring 2016), MS student; Phd student, brigham young univ

hannah reidenbach (spring 2016), BS student;

Tylor O’neill (summer 2015), BS student; wayne state univ

adam ouellette (summer 2014 --- summer 2015), BS student; MD student at Hofstra Univ

Nicholas morse (fall 2014 --- spring 2015), bs student; dmd student univ of detroit mercy

Christian Bonzheim (fall 2014 --- spring 2015), BS student; md student at auc

claire olson (fall 2013 --- summer 2015), bs student, ms program at CMu

Storm Shriver (2014 spring), bs student; senior scientist, Comprehensive lab services

Alec Bergman (2014 spring), bs student;

Mitch Thompson (2013 Fall), bs student;


Dr. Zhao was born in the City of Jilin located in the northeast of China, which is well known for its magnificent natural phenomenon of rime ice in winter. He obtained his B.S. degree in Chemistry from Jilin University and Ph.D. in Bioanalytical Chemistry with Profs. James F. Rusling and John B. Schenkman in the Department of Chemistry at University of Connecticut. During his doctoral studies, Dr. Zhao contributed to designing high-throughput methods for the screening and structural elucidation of DNA-reactive chemicals using electrochemistry and liquid chromatography --- mass spectrometry. After his doctoral training, Dr. Zhao decided to learn more about the biological effect of chemically modified DNA. He was fortunate to have the opportunity to work with Profs. F. Peter Guengerich and Martin Egli at Vanderbilt University School of Medicine. During this time, he obtained solid skills in protein biochemistry, enzymology, nucleic acid chemistry, and macromolecule X-ray crystallography. He elucidated the mutagenic mechanism of N2,3-ethenoguanine, a chemically modified DNA base derived from vinyl chloride exposure and oxidative stress, and concluded that N2,3-ethenoguanine is an important DNA modification for the vinyl chloride-induced human liver carcinoma.

Dr. Zhao started his independent career in the Department of Chemistry and Biochemistry at Central Michigan University in 2013 and relocated to the Department of Chemistry at the University of California, Riverside in 2019. His research program focuses on understanding the chemical and molecular mechanisms of DNA-protein interactions and their biological implications, with an emphasis on DNA replication and repair. He is a dedicated teacher and research mentor. Since 2013, he has mentored 2 postdoctoral fellows, 5 MS students, 2 technicians, and more than 20 undergraduate students. Currently, he is training 5 Ph.D. students, 5 undergraduate students, and 1 postdoctoral fellow. He enjoys working with students at different levels to prepare the next-generation scientists and to cultivate a passion for science.


Linlin Zhao, Ph.D.

MRB 4125

169 Aberdeen Drive

University of California, Riverside

Riverside, CA 92521



Dr. Anal Jana

postdoctoral fellow

current members

Copyright 2013-2022 Zhao Lab | Contact:


Kathleen Urrutia

PhD student

wenxin zhao

phd student

Dr. Prince kumar

postdoctoral fellow

Dr. Wenyan Xu

Research scientist

Yu Hsuan Chen

phd student

Jacob Perkins

phd student

Martin esparza sanchez

Matthew tippin

phd student

seanmory sothy

phd student

Briana hojo

MS student